This paper considers the process of organizational adaptation that took place at the UJA-Federation of New York (UJAFED) between 1990 and 2014. Findings demonstrate that over the last two decades, the UJAFED has operated in turbulent and unstable environments, characterized by a reduction in resources and increasing needs in the communities. These change s have led UJAFED to establish a new domain for its activities that prioritize its mission of Jewish continuity by funding services and programs that emphasize Jewish identity, education, and culture. In this new domain, UJAFED has expanded activities to reach diversiļ¬ed client populations in New York and in Israel. The development of a new domain of activity has had implications for UJAFEDās traditional and new roles, since it has inļ¬uenced its operating strategies, reshaped its mission-driven goals, and changed UJAFEDās relationships with its peer organizations and partners.